生态学理论

• Coordination hypothesis of leaf photosynthesis
• Competitor-stress-ruderal theory
• Leaf Economic Spectrum
• resource co-limitation theory

Coordination hypothesis of leaf photosynthesis

proposed by Chen, Jia-Lin, et al. “Coordination theory of leaf nitrogen distribution in a canopy.” *Oecologia* 93.1 (1993): 63-69. >

大致是说植物光合作用模型不以carbon gain最大化去最优化，而是These two processes are defined as Wc, the Rubisco limited rate of carboxylation, and Wj, the electron transport-limited rate of carboxylation. We suggest that plants allocate nitrogen differentially to, leaves in different canopy layers in such a way that Wc and Wj remain roughly balanced. 

in CLM 5.0, the LUNA model was developed to apply this theory.

https://github.com/ESCOMP/CTSM/blob/059d708bb7ff11ddfe23de73d858f21e021b8bf5/src/biogeophys/LunaMod.F90

https://billsacks.github.io/ctsm-docs/versions/master/html/tech_note/Photosynthetic_Capacity/CLM50_Tech_Note_Photosynthetic_Capacity.html

Competitor-stress-ruderal theory

植物的生态策略主要由三种基本类型组成——竞争型（C）、耐逆型（S）和扰动型（R），它们分别适应不同的资源、压力和干扰环境。

Leaf Economic Spectrum

identified a trade-off between traits related to rapid growth rates (e.g. high leaf nitrogen content, SLA, and photosynthetic rates) and leaf lifespan

resource co-limitation theory

资源共限制理论认为，生物群体的生长和生态过程往往不是由单一资源决定，而是同时受到多个资源的共同限制，例如碳、氮、水分或微量元素的交互作用。 换句话说，生态系统中的生产力和群落结构常常取决于多种资源的协同不足，而不是某一个资源的单独缺乏。

tested by Anu Eskelinena and Susan P. Harrison. “Resource colimitation governs plant community responses to altered precipitation.” *PNAS* 112.42 (2015) 13009-13014

Ecological theory and evidence suggest that plant community biomass and composition may often be jointly controlled by climatic water availability and soil nutrient supply.